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=== Act I ===
{{main|Act I#Synopsis}}
[[Conway]], a truck driver, works as a deliveryman for an antique shop run by a woman named [[Lysette]]. Conway's last job before retirement is to make a delivery to [[5 Dogwood Drive]]. The game opens with a lost Conway, together with his [[Conway's dog|dog]], pulling into the parking lot of a dimly-lit gas station, [[Equus Oils]]. Conway asks for directions, and [[Joseph Wheattree|Joseph]], the owner, informs him that the only way to reach his destination is to take the mysterious [[Route Zero|Route {{Zero}}]]. He tasks Conway to fix the circuit breaker to restore power to the station so that he may use the computer to locate directions. In the basement, Conway encounters [[Emily, Ben, and Bob]] who are playing a game; they ignore him, apparently unable to see or hear him. He finds their lost [[D20|twenty-sided die]] but, on attempting to return it, discovers that they have disappeared. Conway uses the computer to find directions to the [[Márquez Farmhouse]], and also sees an overdue notice from the energy company [[Consolidated Power Company|Consolidated Power Co.]] Joseph tells him that, at the farmhouse, he can talk to [[Weaver Márquez]] who has a better understanding of the roads. As he leaves, he tells Conway that he loaded Weaver's old TV into the back of the truck to take to her.
[[File:Elkhorn Valley.png|thumb|left|''SHANNON: We all lost people down here. Well, not all of us. But most of us.'']]
Back at the farmhouse, Shannon reveals Weaver fled after learning of their family's debt, and tonight was the first time she had seen her since then. Shannon attempts to repair the TV by cleaning moss off the vacuum tubes, and Conway looks in again; this time, however, both the picture on the screen and the barn behind the house begin to warp and separate, forming an image of the opening to the {{Zero}} and the truck driving into it.
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=== Act II ===
{{main|Act II#Synopsis}}
[[File:Bureau.png|thumb|left|''SHANNON: This is weird, but ... do you think we're ''inside'' or ''outside'' right now? <br />CONWAY: Inside. ''/'' Outside. ''/'' Both.'']]
Act II opens with a prelude in which [[Lula Chamberlain]], an installation artist whose work is featured in ''[[Limits & Demonstrations]]'', receives a rejection letter from an [[Gaston Trust for Imagined Architecture|organization]] to which she had submitted her portfolio. She continues on to sort through proposals for reclaiming spaces for alternate to their current function, such as one to reclaim a distillery as a graveyard. If encountered in [[Act I]], [[Carrington]] comes up to her and asks her about the space for his performance.
Thoroughly lost on the {{ZeroLink}}, Conway and Shannon stop at a large, brick, open air building known as the [[Bureau of Reclaimed Spaces]] and ask for directions to [[5 Dogwood Drive]]. Unable to assign them to any of the proper clerks, the receptionist sends them upstairs to Lula who, upon hearing their request, ponders if her former lover [[Joseph]] sent them to her to get her attention or if her former intern [[Weaver]] overestimated her capabilities. Lula suggests the relevant records have not yet been transferred to the new building from the former site of the Bureau, the [[Saint Thomas Church]], and directs them there. She also recommends Conway visit a man named [[Dr. Truman]] to look at his hurt leg.
The two head for the church, which is located inside a [[Random Access Self Storage|self storage facility]], and navigate there by following a series of ritualistic directions. Shannon finds the records while Conway chats with the [[Brandon|janitor]]. Shannon returns with the information but, upon leaving the building, Conway collapses from his injury. Shannon decides their first priority should be to find Dr. Truman and obtain treatment. [[File:Zero.png|thumb|right|''MARY ANN: Just get back on the {{Zero}} and drive until you hit the crystal. Then turn around. It'll make sense once you get on the road.'']]
[[File:Museum roof.png|thumb|left|''EZRA: Julian's my brother. He's a lot bigger and stronger, and he has more feathers than me, but he's still my brother.'']]They return to the Bureau and, while Lula is gone, they find that she has processed the files needed for them to return them to the surface roads. [[Carrington]] appears and asks for a venue recommendation, and Conway is able to suggest either [[Elkhorn Mine]], the self storage facility, or [[Equus Oils]].
Conway and Shannon head above ground and follow Lula's directions to Dr. Truman's house. Upon arriving, however, they find the neighborhood has been torn down and replaced with the [[Museum of Dwellings]]. They talk with many of the residents and meet a young girl named [[Flora]] who tells them Dr. Truman has moved away. On the roof, a young boy named [[Ezra]] tells them he and his brother [[Julian]] fly the museum residents' homes out to the [[forest]] at night which is easier to sleep in, and that Dr. Truman has decided to stay there permanently. Ezra agrees to bring Shannon, Conway, and his dog to the doctor and calls Julian, a giant eagle who snatches them off the roof and flies them to the forest. In the air, the group can observe many situations and locations below, such as the [[Márquez Farmhouse|farmhouse]], gas station, and various other establishments and wandering travelers. [[File:Forest.png|thumb|right|''EZRA: You have a way of falling behind, don't you? Me, I'm always too far ahead. Maybe I should slow down a bit, like you.'']]
The group lands in the forest and walk through the illusory trees as [[Emily, Ben, and Bob]] play a [[Long Journey Home|folk song]] in the foreground. They arrive at Dr. Truman's house and he inspects Conway's leg, who says it's severe but treatable. He gives Conway a dose of an anesthetic called ''Neurypnol TM'', who begins to succumb to the drug. As he falls into into a nebulous sleep, his vision grows black and the walls of the house pull away to reveal the forest beyond.
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=== Act III ===
{{main|Act III#Synopsis}}
Back in the Bureau, the group stands silhouetted in front of Conway's truck, mulling over what's to come next. A tugboat arrives, and as it pulls into view, a horn blares, revealing a gigantic, trumpeting wooly mammoth.
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=== Act IV ===
{{main|Act IV#Synopsis}}